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- protein polyubiquitination
- ubiquitin-protein ligase activity
- protein binding
- nucleus
- nucleolus
- cytoplasm
- multicellular organismal development
- pattern specification process
- zinc ion binding
- protein ubiquitination
- ligase activity
- positive regulation of transforming growth factor beta receptor signaling pathway
- ubiquitin-dependent SMAD protein catabolic process
- positive regulation of protein ubiquitination
- SUMO polymer binding
- protein complex
- positive regulation of transcription, DNA-templated
- SMAD binding
- metal ion binding
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